The training program for a doctoral degree in Molecular Medicine is designed to prepare the student for a career in research and research-related fields.  As such, training activities include of experimental research, coursework, and participation in journal clubs and seminars.  Students entering the Program in Molecular Medicine will conduct original research in one of the laboratories of the Molecular Medicine training faculty.

Courses in one of three emphasis tracks—Cell Biology, Immunobiology, or Medical Biophysics—will provide a foundation in the discipline represented by the chosen track.  Finally, participation in journal clubs and seminars will provide broad, cross-disciplinary exposure to cutting-edge questions and themes related to Molecular Medicine.  In total, the curriculum is designed to train the next generation of forward-thinking and innovative scientists.

The Graduate Program in Molecular Medicine (GPMM) requires 36 credits of coursework (18 or more graded plus 9 credits for the minor, 5 of which are graded) for the Doctorate of Philosophy degree.  All GPMM students will take a core of 8 required units of program core courses. Students will take at least 9 credits of emphasis track courses.   In addition, students will take at least 6 credits of Seminar, 6 credits of Student Seminar, at least 4 credits of Journal Club.  Finally, 18 credits of Dissertation are required. With the approval of the graduate mentor, students may take additional courses for credit.

Below are some useful links:

Graduate Student Handbook

Student Forms

The Core Requirements for all Emphasis Tracks are provided below.  When making course selections, select the appropriate Track using the red navigation buttons below to review specific requirements relating to that that course of study.

Core Courses for all Tracks:

  • CMM/MCB 577: Principles of Cell Biology (Fall)
  • CMM/MCB 695E: Science, Society, & Ethics (Spring)*
  • MCB 580: Introduction to Systems Biology (Fall) OR BIOC 568: Nucleic Acids, Metabolism, and Signaling (Spring)**
  • Student Seminar (CMM 696B or IMB 696B; each Spring and Fall)
  • Journal Club/Colloquium/Work in Progress Seminar (each Spring and Fall). Each Emphasis Track has specific recommendations (see below).

Research Courses for all Tracks:

  • CMM or IMB 900 Research Units - 12 units, 6 per semester until qualifying examshave been successfully completed
  • CMM or IMB 920 Dissertation Units - 18 units, 6 per semester to be started after successful completion of qualifying exams

*Or an equivalent course approved by the GPMM Directors
**Note: the requirement is for students to take either MCB580 or BIOC568, not both

Cell Biology

Cell Biology

To provide students with a foundation for understanding the cellular underpinnings of human health and disease, while advancing the field through hands-on, cutting-edge research.

Medical Biophysics

Medical Biophysics

Apply quantitative multi-disciplinary approaches to molecular and integrative biology with a goal to understand the biophysical basis of health and disease mechanisms, identify drugable targets and contribute to Precision Medicine.



Advance the insights into the rules of engagement that determine the outcome of host-pathogen interactions to develop future therapies for infectious, autoimmune and malignant diseases.